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Autophosphorylation of FGFR1 kinase is mediated by a sequential and precisely ordered reaction
Author(s): Furdui CM, Lew ED, Schlessinger J, Anderson KS
Source: MOLECULAR CELL    Volume: 21    Issue: 5    Pages: 711-717    Published: MAR 3 2006  
Times Cited: 36     References: 29     
Abstract: Tyrosine phosphorylation of cellular proteins induced by extracellular cues serves as a critical mediator in the control of a great variety of cellular processes. Here, we describe an integrated experimental approach including rapid quench methodology and ESI-LC-MS/MS as well as time-resolved ESI-MS to demonstrate that tyrosine autophosphorylation of the catalytic tyrosine kinase domain of FGF-receptor-1 (FGFR1) is mediated by a sequential and precisely ordered reaction. We also demonstrate that the rate of catalysis of two FGFR substrates is enhanced by 50- to 100-fold after autophosphorylation of Y653 in the activation loop, whereas autophosphorylation of the second site in the activation loop (Y654) results in 500- to 1000-fold increase in the rate of substrate phosphorylation. We propose that FGFR1 is activated by a two-step mechanism mediated by strictly ordered and regulated autophosphorylation, suggesting that distinct phosphorylation states may provide both temporal and spatial resolution to receptor signaling.
Document Type: Article
Language: English
Reprint Address: Schlessinger, J (reprint author), Yale Univ, Sch Med, Dept Pharmacol, New Haven, CT 06520 USA
Addresses:
1. Yale Univ, Sch Med, Dept Pharmacol, New Haven, CT 06520 USA
Publisher: CELL PRESS, 1100 MASSACHUSETTS AVE, CAMBRIDGE, MA 02138 USA
Subject Category: Biochemistry & Molecular Biology; Cell Biology
IDS Number: 023OB
ISSN: 1097-2765
DOI: 10.1016/j.molcel.2006.01.022
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